Some minor critiques:
-Rat Rodent at Improved and Greater Rat Race says you can make extra bite attacks per round, but not when. In a full attack? Charge? Whenever you make a bite attack?
-In Inspire Heebie Jeebies, it may be best to allow the player the option of upgrading their Fear effect to Frightened even if they picked Rat Reputation 3+ times (instead of 3 different ways of inflicting Shaken), because area Frighten will be better than having a second way of delivering shaken most of the time.
Otherwise another great work from you, Bhu!